23
Allele
Tests
of
High
Pigment
Genotypes
Using
Root
Anthocyanin
Expression
Jones,
C
.
M
.
1
,
Myers,
J
.
R
.
2
and
Chetelat,
R
.
T
.
1
1
TGRC,
Dept
.
of
Vegetable
Crops,
Univ
.
of
California,
Davis
CA
95616
2
Department
of
Horticulture,
Oregon
State
University,
Corvallis,
OR
97330
Numerous
studies
have
examined
the
carotenoid
content
and
genetics
of
high
pigment
tomatoes
.
The
C
.
M
.
Rick
Tomato
Genetics
Resource
Center
(TGRC)
maintains
accessions
with
hp
-
1,
hp
-
1
w
,
hp
-
2
and
hp
-
2
j
,
as
well
as
dg
and
Ip
.
Historically,
the
nomenclature
of
the
high
pigment
mutants
has
been
somewhat
confusing
.
Thompson
(1955)
originally
proposed
that
an
interaction
of
two
recessive
genes
(
hp
1
and
hp
2
)
was
required
for
the
high
pigment
phenotype
.
Thompson
et
al
(1962)
later
modified
this
to
a
single
gene,
hp
.
Soressi
(1975)
described
hp
-
2,
a
phenotypically
similar
mutation
induced
by
EMS
in
cv
San
Marzano,
which
was
not
an
allele
of
the
original
hp
.
(Note
that
Thompson
(1955)
used
subscript
hp
1
and
hp
2,
and
there
is
no
indication
that
the
orignal
high
pigment
tomato
studied,
Webb
Special,
contains
hp
-
2
.
)
Mochizuki
and
Kamimura
(1986;
1985)
used
seedlings
grown
under
yellow
film
to
test
the
allelism
of
hp,
hp
1,
hp
2
and
dg
and
concluded
they
were
all
mutations
at
the
same
locus
.
The
distinction
between
hp
,
and
hp
1
in
the
Mochizuki
and
Kamimura
(1986;
1985)
study
is
unclear
.
Van
Tuinen
et
al
.
(1997)
suggested
that
hp
-
1
be
used
for
hp
.
Recent
studies
however
have
clarified
that
hp
-
1
and
hp
-
2
are
separate
loci
with
hp
-
1
located
on
chromosome
2
(Yen
et
al
.
,
1997)
and
hp
-
2
located
on
chromosome
1
(Van
Tuinen
et
al
.
,
1997)
.
Kerr
(1965)
described
a
method
using
anthocyanin
expression
in
the
hypocotyl
of
seedlings
below
the
soil
line