APPENDIX

Interim Report of the Committee on Varietal Pedigrees 1995-60

In this second report an attempt has been made to show in part the recent impact of Mendelian and multiple factors on the cultivated tomato. Outside of the mention made to the Spartan Red 8 variety, work with greenhouse types on the American continent has been excluded as it is the hope of the committee, with the help of L. J. Alexander, that in 1961 past greenhouse work will be brought up-to-date in graph form. There follows a list of many of the recent introductions as well as the genetic background of the parents used. Names of the authorities furnishing information follow the varietal names.

Alpine: Powers, Le Roy and Donald H. Scott. U.S.D.A. Horticultural Field Station, Cheyenne, Wyoming (27). See Kenosha variety.

Argentune varieties: See San Marsano paste types.

Australian Tomato types: Notes explainina why it has been found difficult to trace certain Australian tomato varieties. Graham, T. O., Horticulture Department, O.A.C., Guelph, Ontario.

It is known that the Targinnie Red variety came to the United States from Australia, but Dr. C. F. Andrus writes that it may have originated in New Zealand. One possible explanation as to why Australian types are hard to trace and to pin down in that the Yates Seeds Limited of Sydney, Malbourne, Brisbane, and Perth in Australia are associated with allied firms in Tasmania and New Zealand. In the catalogue of this firm in 1954 one can find a list of Chinese types known as Australian Large Red, Tatura Dwarf Globe, Rouge de Marmande, South Australia Early Dwarf Red, and Tatinter. The estalogue states that all Chinese types mentioned have the ability to set flowers and fruit under fairly cool conditions.

While in England, Duncan Hargrave, of the Provincial Horticultural Station, Brooks, Alberta, obtained information from Australia on these Chinese types. He considers that the ability of these to set fruit under cold conditions was segregated out undor growing conditions in China before these types were transferred to Australia. He obtained the Cloche Wonder variety from H. J. Speed & Sons Ltd., Evesham, Worchoster, England. The Cloche Wonder variety he found to be a selection made in England from Tatura Dwarf Globe. The seed of Tatura Dwarf Globe in this came to England from New Zealand (see Globe Trotter variety).

Baby Lea: Darby, L. A., Glasshouse Research Institute, Worthing Road, Little Hampton, Sussex, England. In a latter of April 14, 1959, L. A. Darby states that Baby Lea has been described as a sport out of the variety Potentate. He further states as follows: "I think this is most unlikely.....I am tooting this variety against some of the dwarf alleles including brachytic." Baby Lea has abort internodes and if it is found to be 'br' in type it is the first commercial greenhouse variety to carry the brachytic gene. Baby Lea was introduced in 1946.

Campbell Soup Company 135 and 146 (26). (See Ohio W-R Brookston.)


Targinnie Red  John Baer_______Del Monte  Greater Baltimore
    |                       |                     |
    |                       |___________ Early Baltimore (Mich.)
   M6                              |             1936
U.S.D.A. S._______Rutgers          |
Carolina     |                     |__________________ Red Skin 
             |____Gulf State Market                 |   sp u
              |          y                          |
              |                                     |
              | Sioux_____Burgens   Crack Proof_____Wisconsin 55
              |   u    |  1945 _M.  May trace    |  Partial sp, 
              |        |  in part to             |  tolerant to
              |        |  Crack Proof Pink       |  early and              
              |        |                         |  late blight, 
              |        |                         |  and to 
              |    Glamour                       |  fusarium.
              | Birdseye Snider                  |  Leaf 
              |     1957 u                       |  retantive.
              | Resistance to cracking.          |  Resistant to
S1114         |                                  |  cracking.     
U.S.D.A. S. Carolina_______________ Step 179     |
                      |            Southern Tomato Exchange or
                      |            Kc 54 Campbell Soup Co. Crack
   ___________________|            resistant in Texas. Traces to
   |                               work of Campbell Soup Co., at
   |                               Riverton, N. J.
   |                                   |
Hotset                 ________________|_______
Jacksonville, Texas    |                      |
1957. Resistant to   54-E6______ Ohio W-R     |
heat sterility.      Campbell |  Brookston    |         Improved  
                     Soup Co. |  I 1953       |         Garden
                     Crack    |             54E-14_____ State
                     resistant|             Campbell |    sp
                              |             Soup Co. |
                              |             Crack    |
                              |          Resistant   |
                             146              |      |
                       Campbell Soup Co.     54E14____80
                           1958 ug I                | Campbell
                       Sm (New Jersey, Florida)     | Soup. Co.
                       tolerant to tobacco mosaic.  |
                       Moderate crack resistance.   |
                                                    |
                                                   135 
                                              Campbell Soup Co.
                                                  1958 ug       
                                              Crack resistant.              

Devon Surprise: In the correspondance of the late Dr. J. W. MacArthur at the University, of Toronto, it is noted that Devon Surprise was first located in England as a mutant of the Ailsa Craig variety. The u gens traces back to the Devon Surprise variety and it also posseeses the ad gens for resistance to collar rot.

Early Lethbridge: Nonnecke, I. L. and G. A. Kemp. Research Station, Canadian Department of Agriculture, Lethbridge, Alberta. See R-6-1 variety.

Fireball: See San Marzano paste types.

Firesteel: In a letter of December 17, 1960, S. A. McCrory of the Horticulture Department, Agricultural Experimental Station, Brookings, South Dakota wrote that Clare Barber of Mitchell. South Dakota made the cross and introduced the variety Firesteel. See Sheyenne variety.

Fortuna Original: Firm of Gebr. van de Berg, Naaldwijk, Netherlands, close to 1957 introduced Fortune Original which traces to Moneymaker crossed with Econoom (potato leaf). It carries c and u and is a greenhouse type used in Holland and gaining in importance in Eng (30).

Gem: See New York State varieties (31, 32).

Geneva 11: See New York State varieties (31, 32).

Glamour: Enzie, W. D. Birdseye-Snider. Birdsays Research Laboratory, Albion, N. Y.. See Caupball Soup company varieties 135 and 146.

Globe Trotter and Red Bobs: Hargrave, P. D. , and S. Molnar. Provincial Horticultural Station, Brooks, Alberta (13).

                                     Tatura Dwarf Globe
                                           Australia
                                            sp, nt
                                        Cold tolerant
                                              |
                                              |
                                              |
                                              |
All Red_________________________________ Cloche Wonder
sp u                 |                    England sp
                     |           Selection made for cloche work
                     |                  Cold tolerant
          ___________|______________________
          |                                | 
Globe Trotter                             Red Bobs
1960 sp, u                                 1960 sp
                                     Blemish Resistant.
                                 Resistant to cold sterility.

Hotset: Young, P. A., Tomato Disease Laboratory. Agricultural Experimental Station, Jacksonville, Texas (3). See Campbell Soup Company varieties 135 and 146.

Juno: In a letter of May 23, 1959, Ole Frederiksen, L. Daehnfeld Ltd., Odense, Denmark wrote that he could not trace the variety Immun in Prior Beta which is cold tolerant and possesses the potato leaf character. He was procuring seed of this variety from Czechoslovakia. He had crossed Immun Prior Beta II vith Moneymaker and this resulted in an extra early Danish, open ground, variety named Juno.

Kenosha: Foskett, R. L. Departnmt of Horticulture, State University. Fort Collins, Colorado (10).

Denmark u____________________________________L. pimpinellifolium
Tolerant to late blight                  |
                                         |                                         
Denmark_________________________________ F1
                                  |
                                  |
                                  F6____________ Early Baltimore
                                     |   
                                     |
Loran  Blood (Utah)__________________|____Alpine (U.S.D.A. Wyo.)
1952   Ve             |                           1949
                      |                   Probably cold tolerant
                      |                     Altitude 6200 feet
                   Kenosha
                     1958
                 Altitude 5100

Loran Blood: See 'Sources of Verticilium Wilt Resistance on Tomato Variatis'.

Marion: Epps, William, and W. R. Sitterly. Clemson College, Agricultural experimental Station. Charleston, South Carolina (9).

L. pimpinellifolium___________ Marglobe
Sm (S. Carolina)           |
                           |
Pan American              SC 35
I (U.S.D.A. Beltsville)___Sm (S. Carolina)
                        |
                        |
                        |
                       SC 121___________ Pan American
                 Sm (S. Carolina) |         I
                                  |
                                  |
                                SC 137___________ Step 89 (1950)
                            Sm (s. Carolina)  | Became Homestead 
                                              | variety 1952 I 
                                              | Semi-determinate
                                              |
                                Tolerant to early blight
                                Partial crack resistance
                                Foliage cover and retention
                                equal to Rutgers and superior
                                to Homestead.

Missouri S-34: Tucker, C. M., Horticulture Department, University of Missouri, Columbia (19, 20).

              Earliana_________________ L. pimpinellifolium
                                   |    I (Missouri 160)
          Break O'Day______________|    PI 79532
                               |
       Bonny Best______________| 
                           |
   Break O'Day_____________|
                       |
Rutgers________________|
                 |
      ___________|___________
      |                     |
Missouri S-34             Mosage
1951 I                    1958

MO Surprise F1 Hybrid: Lambeth, Victor H. Horticulture Department, University of Missouri, Columbia. See Mozark variety (20). Mosage: Lambeth, Victor H. Horticulture Department, University of Missouri, Columbia. See Mozark variety (20).

Mosage: Lambeth, Victor H. Horticulture Department. University of Missouri, Columbia. Missouri S-34 variety (19).

Moscow: The following information was suppllied by Dr. Orson Cannon. In his letter of January 18, 1961 he states: "..... the following, which was related to me by the late Pat Winters, probably is correct. He said that a Japanese grower in Utah by the nme of Masuka selected an off-type plant in a field of Marglobe. Its offspring did so well in the area that the variety was adopted by many of the growers. Gradually the name was changed from Masuka to Moscow. Supporting this information is the fact that Moscow was grown commercially in Utah before it was grown elsewhere. When I came to Utah in 1948 the variety was being grown in about 75% of the acreage of the state, and many gravers had their own strains, each of which was reputed to be better than the strains belonging to others."

Mozark: Pedigree is listed in the following publication: Lambeth. Victor H. 1958. The Mozark tomato. Missouri A.E.S. Res. Bull. 680, 1-8. The MO Surprize F1 Hybrid is a cross between Mozark I sp u and Sioux u and it was introduced by Victor H. Lambeth in 1959 (20).

Nagcarlang Tomato type from the Philippines. Graham, T. O., Horticulture Department, O.A.C. Guelph, Ontario.

The description of the Nagcarlang tomato type which follows has been made up in part from information given by Dr. Eugenio E. Cruz, Director, Bureau of Plant Industry. Manila, Republic of the Philippines and from Dr. Howard Peto, Peto Seed Company, Saticoy, California. The Nagcarlang tomato is the onlytype which will set fruit in high humidity areas throughout the Philippines. For this reason, during the winter of 1948--49, Dr. Peto called at Manila to see if the Nagcarlang tomato type could be used in a breeding programs so that a canning tomato for the Philippines would result. He also took tomato varieties, over to Mindanao and the Southern Philippines,of which Kolea and Anahu have proven adaptable.

Under Ontario greenhouse condition the flowers of Nagearlang are open in type and the pistils are exposed. As a result one must place pollen on the pistils if pollination is to take place. In the tropics high insect activity possibly carries out cross polination.

The Nagcarlang type has come into Ontario in two shipments. The first seed obtained has been written up in Scientific Agriculture 32:57-66, 1952. As far as is known the only surviving member in breeding programmes of the first shipment made to Ontario in 1948 is the type known as Philippine #2. Dr. Eugenio E. Cruz forwarded Nagcarlang to Guelph in the second shipment made in 1959, this seed having been gathered at Laguna in the Philippines.

In the Philippines Nagcarlang is known to be a wild-type tomato. Its history is unknown. It is not a variety as no two plants are alike. It is simply a heterozygous form which has survived in the jungles of the Philippines and which has also been brought under cultivation and even sold commercially. Dr, Peto has verbally stated that in the Philippines the Nagcarlang type has survived-and become pollinated in areas with 100 inches of rainfall per season. He stated that it ia hard to believe that such wild-type fruit could be sold commercially. However, he saw Nagcarlang fruits being sold when he visited the open market in Manila. He noted that the flowers hang down, shed the vater, and do not experience as much blossom drop rainy weather aa is experienced by commercial types.

While no two Nagcarlang plants are the same there are points in common. For example, practically all Nagcarlang fruits are small, very rough. and with a purplish "muddy" fruit colour. The fruits are firm and under Ontario conditions flat and sweet in flavour. They mostly ripen late in the season. The plants are very robust, and dense with healthy foliage. While Dr. Peto attributes fruit set under humid conditions to the flowers, the fact that the pollen remains active may have a chemical basis as the plants and fruits of the Nagcarlang type have a distinct purplish colour.

In the Philipyines the Nagcarlang tomato is resistant to disease and to periodic drought. In Ontario it is tolerant to late blight. Under Ontario conditions it will set fruit at temperatures as low as 45 deg. F, which is 10 degrees below the point at which tomatoes normally set fruit. In other words, it is cold tolerant, It is interesting to note that in the tropics Nagcarlang is resistant to sterility caused by damp conditions and in Ontario, far removed from-the tropics, it is resistant to sterility caused by cool conditions.

In the tropics the point of interest in to see if, through a breeding programs, the bridge can be gapped between the wild Nagcarlang type and the commercial varieties found on the North American continent. This would allow the commercial canning of tomatoes to be carried out in certain tropical areas such as the Philippines, where high humidity makes this difficult at the present time.

New York State Varieties: Gem, Geneva 11, Red Jacket. Tapley, W. T., Agricultural Experimental Station, Geneva, New York State (31, 32).



Stokesdale__________ Bounty__________ Fruit from an unknown
              |       sp u     |      variety displayed at
              |                |      Palmyra County Fair,
              |                |      N. Y.
              |                |
             Gem           Red Jacket_____ Line resistant to
           1947 sp u       1947 c u    |   Verticillium wilt, Ve
                                       |
                                    Geneva 11
                                    1957 c Ve
                             Late blight resistance

For an explanation as to the monogenic and multigenic resistance displayed by Geneva 11 to late blight see Galleghy M. E. 1960, Resistance to the Late Blight Fungus in Tomato, Proc. Science Seminar. Campbell Soup Company, p. 113-135.

Ohio W-R Brookston: Pedigree forwarded by latter on January 6, 1961, and variety is described by Alexander, L. J. 1954. Ohio Farm and Home Research, Jan. - Feb., p. 8. Variety was introduced by Dr. L. J. Alexander of the Department of Botany and Plant Pathology, Agricultural Experimental Station, Wooster, Ohio.

Bonny Best^2___________________ L. pimpinelifolium
                    |           I  (Missouri 160)
                    |           P.I. 79532
                    |
                    |__________ Bonny Best
                       |
                       |__________ Bonny Best
                          |
                          |___________ Break O'Day
                             |
                            F2____________ Rutgers
                                |
                               F2_____________ Rutgers
                                      |  
                                      |
                                     F5
                                      |
                                      |
                             Ohio W-R Brookston
                                   I 1953

Ontario variety: In a letter of March 8, 1960, R. D. Peel wrote that the Campbell Soup Company introduced the Ontario variety in 1946 and that this even ripening type was obtained from a cross between Rutgers and Bounty.

Owyhee: Simpson, W. R. Branch Experimental Station of the University of Idaho, Parma, Idaho (29).

L. peruvianum_______ Bounty sp      Red river________Coopers
Curly top       |                               |    Special
resistant       |                               |      sp
selection       |____________________________Bison sp
                                       |
Stokesdale_____ All Red sp u         Self
            |                          |
         Sioux u____________________ Curly Top Resistant Selection
            |                     |
            |                   Self 
            |                     |
            |__________________ Curly Top Resistant  Selection
            |                |                               
            |              Self
            |                |
            |_____________ Curly Top Resistant Selection
            |           |
            |         Self
            |           |
            |________ Curly Top Resistant Selection
                   |
                 Self
                   |
                 Self
                   |
                 Owyhee
                 1959 sp
                 Curly Top Resistant
Complex mode of inheritance of resistance to Curly Top is as yet not known.

Pearson B (Pearson XL), Pearson VF6, and Pearson VF 11: See "Sources of Verticillium Wilt Resistance on Tomato Varieties."

Peru Wild: Lesley, J. W. University of California, Citrus Experimental Station, Riverside, California (8, 21). See also 'Sources of Verticillium Wilt Resistance on Tomato Varieties'.

Peru Wild was collected by the late O. F. Cook of the U.S.D.A. As far as is known it did not carry an S.P.I. number. It was obtained from Dr. Cook by J. W. Lesley and placed under California accession number 467 it was found susceptible to Curly Top (Hilgardia 6(2) 1930).

Several lines were later sent to Loran Blood of the U.S.D.A. at Logan, Utah, by J. W. Lesley, as it was considered that amongst these a line might be located with resistance to Verticillium Wilt. Within these lines Loran Blood located resistance in the Peru Wild type, and he placed this line under number as Utah 665.

Utah 665 is not the same as 52.68.13 from the Citrus Experimental Station, which is also resistant to Verticillium, and comes from a cross made by F. O. Holmes of L. esculentum x L. chilense (2). As far as is known no comercial varieties have had this last mentioned type as a parent (5).

Quebec varieties introduced under a Joint Programme of the Agriculture for the Province of Quebec and Laval University, Qeebec city, P. Q. (34).

Asgrow Scarlet Dawn______ Italian Red Pear______ Chatam
       1934           |         Italy         |  1950 sp u
                      |   Seed received 1938  |
                      |                       |
King______ Ponderosa  |                   Quebec 152     
Humbert |    1892     |                   1950
paste   |             |                   Cold tolerant
type    |             |
Italy   |             |   
        |             |
    Nystate___________F1
  Geneva, N.Y. |      |
    1935       |      |
               |      |
           Quebec 13  |
           1947       |
           Staking    F8
           type       |
                      |
                   Quebec 5____________________ Red Cloud
                    1946             |          1944 sp u
                Staking type         |
                Resistant to         |
                  cracking           |
                                     |
                              _______|_______
                             /               \
                        Quebec 309        Quebec 314  
                        1954 sp u         1954 sp
                                          Cold tolerant

R-6-1: Calberg, C. W., Research Station, Canadian Department of Agriculture, Swift Current, Saskatchewan (4, 18).

Bounty__________ Farthest Nort_____________ Polar Circle
sp u       |         sp                 |
           |                        ____|____   
           |                       /         \
Redskin___F-2-5           Early Lethbridge   Earlinorth
sp u    | Swift Current   1953 sp u. Pale    1952 sp u
        |                 green u-type.      White green u-type.
        |                 Resistant to cold  Resistant to cold
      R-6-1               sterility.         sterility.
   1957 sp u              Altitude 3020 feet.
   Resistant to
   cold sterility.

Red Global: Horticultural Research Department, Experimental Station, Baton Rouge, Louisiana. Miller, J. C., T. P. Hernandez, and E. C. Tims (22).

Gulf State Market y
        |
        |       
Louisiana Slicer y_______________________ Marglobe
 Mutation 1938         |
Julian C. Miller       |
                       |
                       F1____________________ Marglobe
                    Louisiana    |
                                 |
                                 |
                             Red Global
                         1959 I (Louisiana)
                    Foliage retention in summer and
                   fall. Resistance to heat sterility.

Red-Jacket: See New York State varieties (31, 32).

Roma: Porte, W. S., U.S.D.A. Agricultural Research Service, Beltsville, Maryland (25). See San Marzano paste types.

Red Top: Tapley, W. T., Horticulture Department, Agricultural Experimental, Station, Geneva, N. Y. (33). See San Marzano paste types.

Red Top V9 or VR No.9: G. C. Hanna. Vegetable Crops Department, Agricultural Experimental Station, Davis California (6). See San Marzano paste types.

San Marzano paste types: While in the Argentine, Dr. Abelardo Piovano, in his article Variedades Comerciales de Tomates, B.A.P. Resista Agricola, April 1932, states that the King Humbert paste tomato is an old variety and that it originated with Fratelli-Ingegnoli, Milano, Italy. He also states that the paste types San Marzano and Vittorio Emanele are superior to the King Hubert variety.

In publication 3, for the period 1951-53 of the Facultad de Ciencias Agrarias, Universidad Nacional de Cuyo, Argentine, Dr. Piovano and Luis O.

Melis state, on page 5, that San Marzano has broken into various lines, namely, Pero 20, Santa Catalina, Pera Amarilla, etc. In 1951 the Joseph Harris Company of New York State lists the paste type Italian Red Pear (Pomo d'oro a Pera). This could be another selection out of San Marzano.

The chances are that, in areas outside Italy, unless selection is kept up, these paste types tend to produce small-sized fruit and that many of the known selections catalouged in the past reflect periods at which San Marzano was rogued and brought back into line.

Recent breeding of paste types has led to the introduction of the tomato variety Fireball, and the paste types Roma, Red Top V9, and San Pablo. These four varieties as well as other types are traced in the two pedigrees which follow.

San Marzano is also being used in breeding programmes in South Carolina Florida, and Texas. In 1960 A. L. Harrison has reported in his article 'Breeding of Disease Resistance to Nematode', in Proceedings Plant Science Seminar - 1960, Campbell Soup Company, page 65 as follows:

'While the writer was in the Florida, some of this material now backcrossed to San Marzano, was obtained from C. D. Andrus of the Southeastern Vegetable Breeding Laboratory in Charleston, South Carolina. All of the Texas root knot resistant lines are descendants from this cross, (L. esculentum var. Michingan State Forcing x L. peruvanum) x San Marzano'.

A. San Marzano paste types from United States as well as the non-paste variety, Fireball (6,33).

Marglobe_____ L. pimpinellifolium    Red Top Paste
          |   I (Missouri 160)       Geneva N. Y.
          |   P. I. 79532            1952 sp u o
          |                          Possible parentage
          |                          Italian Red Pear o
          |                                |
     Pan American____________ San Marzano  |
     I (U.S.D.A.           |  Paste o      |
       Beltsville)         |               |
                           |               |      
                    Paste Selection___ Red Top_____ Early small-
                         I o        |  sp u o    |  vined
                                    |  Resistant |  Californian.
                                    |  to        |  Verticillium
                                    |  cracking  |  resistant
                                    |      |     |  type Ve
                                    |      |     |
                               Roma paste  |  Red Top V9
                               1955 I sp o |  1959 Ve sp u 
                               (U.S.D.A.   |  (California)
                               Beltsville) |      
                                           |
Victor (Mich.)__________________________ Red Top
  sp u                     |             Lineclub 86
                           |             Agr. Exper. Stn.
                           |             Geneva, N. Y.
                 Fireball (Earlibird)    sp u o
                 Harris Seed Co.
                 1952 sp u
                 Resistant to cold sterility.            


B. San Marzano paste types from the Argentine as well as several other nonpaste types from the same source.

There follows the pedigree of the varieties introduced by Professor Abelardo Piovano formerly of the National University of the Argentine and now employed in Italy with Cirio, San Giovanni A Teduccio, Napoli. The varieties which trace to Professor Piovano are listed as from the Argentine. The Perfection variety, which is one of the original parents in the Argentinian programe, came from the U.S.A. It traces back to the Livingston Seed Company in 1880 and was located in a field of the Acme variety first listed by Livingston in 1875 (23, 24).

Marmande______ Perfection_____________ Marvilla Del Mercado
L. Clause  |     U.S.A.        |       (Marvel of the Market)
France     |      1880         |       L. Clause, Bretigny,
1930 sp    |                   |       France.
           |                   |
        Marman            Mandocina_________ San Marzano paste
    Argentine 1939           Argentine  |    selection Piovano o
   Semi-determinate            1934     |    Argentine
           |                   |        |
           |                   | San Marzano Chico_____ Bounty
           |                   |  Argentine 1946    |  N. Dakota
           |                   |        o           |  sp u
           |                   |                    |
      Marman Liso              |                    |
    Argentine 1949_________ Mendocina               |
           |                   |             Cirio 49 paste
           |                   |             Argentine 1950
         Cuyano             J. Peron              o sp
       Argentine         Argentine 1948             |
         1952            Resistant to               |
                         disease                    |
                               |                    |
                          Mendoza 44         San Pablo paste
                          Argentine 1952     Argentine 1958
                               |                 o sp
                      _________|________            
                     /                  \
                 3-670                  3-672
                 Argentine 1958         Argentine 1958

Sheyenne: Holland, Neal J. and E. P. Lana. Horticulture Department, State College Station, Fargo, North Dakota (15).

Bounty_______ Stokesdale     Bison________ Pritchard
sp u    |                      sp     |        sp
        |                             |
        |                        Firesteel______ Morden Manitoba   
        |                        S. Dakota   |     Selection
        |                        1935 sp     |        sp
        |____________________________________|
                           |
                           |
                       Sheyenne
                       1959 sp u
                   Blemish resistant.
                   Foliage retention.

Sources of Verticillium Wilt Resistance in Tomato Varieties: Orson S. Cannon, Head, Department of Botany and Plant Pathology, Utah State University. Formerly Plant Pathologist, U.S.D.A. Agricultural Research Service, Logan, Utah.

The verticillium wilt resistant tomatoes of today go back to Utah No. 665 (Peru Wild) for their resistance, and they come from one of two sources: (1) H. L. Blood gave some of his breeding stocks to the late Pat Winters of California Packine Corporation who did breeding work in Utah and California. A number of the present day verticillium wilt resistant varieties have come from his stocks. (2) All of the other verticillium wilt resistant varieties either were developed at the Utah Station or have resulted from crosses in which Utah varieties served as the original source of resistance. These statements, of course, leave the varieties Riverside, Essar, and Simi out of consideration because they do not have the type of resistance reported by Schaible, Cannon and Waddoups (28).

Peru Wild (Utah 665) was part of a collection of tmato seed received by H. L. Blood from J. W. Lesley in 1932. According to Dr. Blood's records, Lesley had previously received this tomato from O. F. Cook. In 1948 Dr. Lesley did not have seed of Peru Wild on hand. Dr. Blood died in 1948 and he left several selections of No. 665. All of them were cherry type tomatoes and all of them were segregating for resistance to verticillium wilt. Viable seed of No. 665 is still in the Utah collection, which is now under the direction of Mark W. Martin.

The original work on the varieties Loran Blood, VR Moscov, Pearson VF6, Pearson VF11, Pearison B and Pearson XL was done at the Utah Agricultural Experiment Station, as outlined below, while the author was employed by the Agricultural. Research Service of the United States Department of Agriculture.

Loran Blood Ve = (Peru Wild x one or more other varieties) Stone In a memorandum dated January 14, 1952 seed of VR No. 4 was made available for testing and for increase of seed. VR No. 4 was released as Loran Blood in Decmber 1952 (7). Pat Winters of California Packing Corporation received some advanced breeding stocks from Dr. Blood. He crossed one of these with Stone. Winnters and the author selected a plant from an F2 population of this cross in 1948. The variety Loran Blood resulted from selections among its progeny.

VR Moscow Ve sp = W6 x Moscow^3. W6 = Peru Wild x Century VR No. 11 of the January 14, 1952 memorandum became VR Moscow upon release (7). The present stocks of VR Moscow are W6 x Moscow^5 (sometimes referred to as M82).

Pearson VF6 and Pearson VF11 Ve I sp = (59-1 x Southland) Pearson^7. 59-1 + ((Peru Wild x Century) Bonny Best) Moscow

All of the crossing work and testing for resistance was done at Logan, Utah by the author. The F2 of the last backcross was grown at Davis, California by G. C. Hanna, and selections in the F2 and F3 were made together by Hanna and the author. The varieties were released 1959 (6).

Pearson B and Pearson XL Ve sp = 047 = ((Peru Wild x Century) Victor) Pearson^3 These two are the same variety. On March 2, 1953 the author sent seed of No. 047, a Pearson type tomato resistant to Verticillium wilt, to Dr. J. W.

Lesley for trial. Dr. Lesley had plants grown from this seed by Clarence R. Brown, Wholesale Seed Grower, San Juan Capistrano, California. Mr. Brown selected seed from some of these plants, and their offspring later were released by him to seedsmen as Pearson B and Pearson XL.

Spartan Red 8: Homma Shigemi. Department of Horticulture, Agricultural Experimental Station, East Lansing, Michigan (1, 14, 16).

                                      Earliana_____ L. pimpine-
                                                 |     llifolium
                                                 |  I (Missouri 160)
                                                 |  P.I. 79532
                                  Break O'Day____|
                                               |
                                Ponderosa______|
                                             |
                              Greater        |
                              Baltimore______|
                                           |
                                           |
Porter______Bounty   Livingston Globe_____ M-8-2
Possibly  |   sp u        y            |   I (Ohio) 
Resistant |                            |
to heat   |      Association Globe_____|
sterility |            y             | 
          |                          |
          |   Selection A Globe______|
          |        Ohio y          |     
Nebraska 12____________________ Ohio WR Globe
   u                   |        1948 I y 
                       |        Greenhouse type.
                       |
               Spartan Red 8 1960 u
          Resistant to heat sterility
                 Greenhouse type.

Stair: Stevenson, E. C., K. W. Johnson, and M. L. Tomes. Department of Horticulture and Botany and Plant Pathology, Purdue University Agricultural Experimental Station, Lafayette, Indiana (17).

Pan American_________________________________ Indiana Baltimore
I (U.S.D.A. Beltsville)            |               1934
                                   |
                            Indiana 1366____________ Rutgers
                                 I            |      Tolerant to
                                              |      fusarium
                                              | 
                                            Stair
                                            1958 I

Stick: Young, P. A., Tomato Disease Laboratory, Agricultural Experimental Station, Jacksonville, Texas. In Jour. of Hered. 46:243-244, 1955, P. A. Young describes a dominant mutant which he termed Cu or Carl and which he located in 1953 as a mutant on a plant of the Stokesdale variety. Because the vines are distinctly non-suckering in type, Gleckers Seedsmen of Metamora, Ohio, listed Cu in 1959 as Stick. This mutant has stripes on the fruit similar in appearance to the gs factor.

Tanggula: Graham, T. O., Horticulture Department, O.A.C., Guelph, Ontario (12, 36).

Earliana
   |
   |
Striped Mutant________________________________ Harrow
Possible allele                  |             sp u
of gs                            |             Blemish resistant
                                 |
                              Tanggula
                              1960 sp u
                          Blemish resistant

Verticillium Wilt resistant varieties: See 'Peru Wild'. See also 'Sources of Verticillium Wilt Resistance on Tomato Varieties'.

VR Moscow: See 'Sources of Verticillium Wilt Resistance on Tomato Varieties'. Wisconsin 55: Gabelman, W. H. and G. C. Pound, Department of Horticulture, College of Agriculture, Madison 6, Wisconsin (11, 35). See Campbell Soup Company varieties 135 and 146.

Literature Cited

1. Alexander, L. J. 1949. Ohio W. R. Globe. Ohio A.E.S. Res. Bull. 689.

2. Alexander, L. J., and M. M. Hoover. 1955. Disease Resistance in Wild Species of Tomato. Ohio A.E.S. Res. Bull. 752, p. 72.

3. Andrus, C. F., and G. B. Reynard. 1945. Resistance to Septoria Leaf Spot and its Inheritance in Tomatoes. Phytopathology 35:16-24.

4. Calberg, C. W. 1959. (Research Sta., Swift Current., Sask.) Correspondence.

5. Cannon, O. S. 1960. (Utah State Univ.). Correspondence.

6. Cannon, O. S., and G. C. Hanna. 1959. New Disease Resistant Tomatoes. Calif. Agr. 3(3):7.

7. Cannon, Orson S., and Victor Waddoups. 1952. Loran Blood and V. R. Moscow, Two New Varticillium Wilt Resistant Tomatoes for Utah. Farm and Home Science (Utah) 13:74-75.

8. Doolittle, S. P. 1954. The Use of Wild Lycopersicon Species for Tomato Disease Control. Phytopathology 44:409.

9. Epps, William, and W. R. Sitterly. 1958-59. S. Carolina Clemson Collage .S. 72nd Annual Rept., p. 3.

10. Foskett, R. L. 1960. (Univ. Colorado). Correspondence.

11. Gabelman, W. H., and G. C. Pound. 1960. (Univ. Wisconsin). Correspondence.

12. Graham, T. O., E. W. Franklin, A. Zitnak and B. Barabas. 1958-59. Progress Report, Vegetables for Processing, p. 35.

13. Hargrave, P. D. 1959. The Brooks Bulletin. Prov. Hort. Sta., Brooks, Alta., p. l.

14. Hoffman, I. C. 1940. Progress in Greenhouse Vegetable Breeding. Proc. Ohio Veg., and Potato Growers Assoc. 25th Annual Rept., p. 103-106.

15. Holland, Neal S., and E. P. Lana. 1959. The Sheyenne Tomato. N. Dakota Farm Res. Bi-monthly Bull. 21(2):16-18.

16. Honma, Shigemi. 1960. Spartan Red 8. Mich. A.E.S. one page announcement of release.

17. Johnson, K. W. 1959. (Pardue Univ.). Correspondence.

18. Kemp, G. A., and I. L. Nonnecke. 1960. Difference in Intensity of Unripe Fruit Colour in Tomato. Can. J. Plant Sci. 40:306-309

19. Lambeth, Victor H. 1959. The Mosage Tomato, Missouri A.E.S. Res. Bull. 681.

20. Lambeth, Victor E. 1959-60. (Univ. Missouri). Correspondence.

21. Lesley, J. W. 1960. (Univ. CAlifornia Citrus Exper. Sta.). Correspondence.

22. Miller, Julian C. 1960. (Univ. Louisiana). Correspondence.

23. Morrison, Gordon. 1938. Tomato Varieties. Mich. A.E.S. Bull. 29, p. 15 and 49.

24. Piovano, Abelardo. 1960. (National Univ., Argentine). Correspondence.

25. Porte, W. S. 1960. (U.S.D.A. Agr. Res. Service, Beltsville, Maryland). Correspondence.

26. Reynard, G. B. 1960. Breeding Tomatoes for Resistance to Fruit Cracking. Proc. Plant Science Seminar. Campbell Soup Co. p. 93-108.

27. Salter, R. M. 1949. Memorandum to Seed Companies and Vegetable Growers Relative to the Release of an Inbred Line of Tomato for use as a Parent in Production of Tomato Hybrids. U.S.D.A. Agr. Res. Admin. Bureau of Plant Industry, Soils, and Agr. Engineering, Jan. 18.

28. Schaible, Lester, Orson S. Cannon, and Victor Waddoups. 1951 Inheritance of Resistance to Verticillium Wilt in a Tomato Cross. Phytopathology 41:986-990.

29. Simpson, W. R. 1959. The Owyhe Tomato. Idaho's Curly-Top Resistant Variety. Idaho A.B.S. Bull. 298.

30. Szteyn, K. 1959. (Institute of Hort. Plant Breeding, Wageningen, Holland). Correspondence.

31. Tapley, W. T. 1947. New Tomato Varieties Ready for Wider Tests. Farm Res. 13(1):7.

32. Taplay, W. T. 1947. Sixty-sixth Annual Rept., Geneva A.E.S., p. 53.

33. Taplay, W. T. 1952. A New Paste Tomato. Farm Res. 18(1O):3.

34. Vandal, J. O. 1958-59. (Univ. Laval). Correspondence.

35. Walker, J. C., G. S. Pound, and J. E. Kuntz. 1948. Development of Wisconsin 55 Tomato. Univ. Wis. A.E.S. Bull. 478.

36. Young P. A., and J. W. MacArthur. 1947. Horticultural Characters of Tomatoes. Texas A.E.S. Bull. 698, p. 34.

Committee on Varietal Pedigrees

Alexander, L. J.         Hernandez, T. P.       Marx, G. A.
Darby,  L.  A.           Homa, S.               Miller, J. C.
Fookett, R. W.           John, C. A. (pending)  Odland, M. L.
Frazer,  W. A.           Johnson, K. W.         Peto, H.
Gabelman, W. H.          Kemp, G. A.            Stark, F. C.
Gilbert, J. C.           Lambeth, V. N.         Tomes, M. L.
Graham, T. O., Chairman  Lana, E. P.            Young, P. A.

It is considered that the committee would be further strengthened by securing a member each from Australia, France, Italy, and Japan. Full co-operation is assured from Holland. A member is also needed from Florida.

Background of Certain Varieties Needed

The parentage and genetic background of the varieties is needed: Century, Del Monte, and Porter.